Opendata, web and dolomites


The earliest migration of Homo sapiens in Southern Europe: understanding the biocultural processes that define our uniqueness

Total Cost €


EC-Contrib. €






 SUCCESS project word cloud

Explore the words cloud of the SUCCESS project. It provides you a very rough idea of what is the project "SUCCESS" about.

replaced    world    independently    bone    occurred    45    understand    adaptive    documents    hotly    pattern    biocultural    cutting    cultures    dating    questions    bp    paleoanthropology    lithic    plays    species    question    replacement    final    transitional    autochthons    prehistory    technocomplexes    arrived    ca    generally    implies    rest    variability    arrival    mousterian    extinction    ago    africa    paleolithic    favoured    edge    modeling    timing    tools    italy    ecological    anatomically    context    position    ornaments    zooarchaeological    amh    modern    human    cal    interdisciplinary    hominin    upper    debated    calendar    survey    shifts    middle    successful    pigments    dramatic    isotope    australia    protocols    multidisciplinary    35    paleoanthropological    archaeological    asia    transition    period    revolves    arrivals    paleoecological    radiated    biological    neandertals    cultural    humans    statistical    amhs    neandertal    suggests    pivotal    sites    geographic    southern    investigations    demise    appearance   

Project "SUCCESS" data sheet

The following table provides information about the project.


Organization address
address: VIA ZAMBONI 33
postcode: 40126

contact info
title: n.a.
name: n.a.
surname: n.a.
function: n.a.
email: n.a.
telephone: n.a.
fax: n.a.

 Coordinator Country Italy [IT]
 Project website
 Total cost 1˙993˙811 €
 EC max contribution 1˙993˙811 € (100%)
 Programme 1. H2020-EU.1.1. (EXCELLENT SCIENCE - European Research Council (ERC))
 Code Call ERC-2016-COG
 Funding Scheme ERC-COG
 Starting year 2017
 Duration (year-month-day) from 2017-05-01   to  2022-04-30


Take a look of project's partnership.

# participants  country  role  EC contrib. [€] 


 Project objective

Anatomically modern humans (AMHs) radiated out of Africa into the rest of the world around 60,000-50,000 years ago. While various evidence suggests arrivals of AMHs in Asia and Australia ca. 45,000 calendar years before present (cal BP), the timing and pattern of the biological and cultural shifts that occurred in Europe around 50,000 to 35,000 cal BP, are hotly debated and are considered to be among the most important questions in paleoanthropology. This period documents 1) the demise of the autochthons Neandertals and their replacement by AMHs as well as 2) dramatic changes in human behaviour and the appearance of various technocomplexes that replaced pre-existing Mousterian cultures. Debate revolves around the question of whether modern behaviour was independently achieved by Neandertals or is directly related to the arrival of AMH. In this context Italy plays a pivotal role due to its geographic position and ecological variability, and key archaeological sites with transitional and early Upper Paleolithic cultures. This interdisciplinary and multidisciplinary project aims to understand when AMHs arrived in Southern Europe, the biocultural processes that favoured their successful adaptation and the final cause of Neandertal extinction. The project involves survey in Italy and field investigations of key archaeological sites dating to the Middle-to-Upper Paleolithic Transition, paleoecological, zooarchaeological, paleoanthropological and isotope analyses, studies of pigments, ornaments, lithic and bone tools, as well as statistical modeling using cutting-edge protocols. The results of this work will be of pivotal importance for understanding a key period in European prehistory and, more generally, the biocultural, adaptive and ecological characteristics that make our species successful and unique. It implies that this project will be crucial for understanding the replacement of all hominin species by AMHs around the world.


year authors and title journal last update
List of publications.
2019 Rita Sorrentino, Nicholas B. Stephens, Kristian J. Carlson, Carla Figus, Luca Fiorenza, Stephen Frost, William Harcourt‐Smith, William Parr, Jaap Saers, Kevin Turley, Stephen Wroe, Maria G. Belcastro, Timothy M. Ryan, Stefano Benazzi
The influence of mobility strategy on the modern human talus
published pages: 456-469, ISSN: 0002-9483, DOI: 10.1002/ajpa.23976
American Journal of Physical Anthropology 171/3 2020-03-05
2019 Simona Arrighi, Adriana Moroni, Laura Tassoni, Francesco Boschin, Federica Badino, Eugenio Bortolini, Paolo Boscato, Jacopo Crezzini, Carla Figus, Manuela Forte, Federico Lugli, Giulia Marciani, Gregorio Oxilia, Fabio Negrino, Julien Riel-Salvatore, Matteo Romandini, Marco Peresani, Enza Elena Spinapolice, Annamaria Ronchitelli, Stefano Benazzi
Bone tools, ornaments and other unusual objects during the Middle to Upper Palaeolithic transition in Italy
published pages: , ISSN: 1040-6182, DOI: 10.1016/j.quaint.2019.11.016
Quaternary International 2020-03-05
2020 María Dolores Garralda, Bruno Maureille, Adeline Le Cabec, Gregorio Oxilia, Stefano Benazzi, Matthew M. Skinner, Jean-Jacques Hublin, Bernard Vandermeersch
The Neanderthal teeth from Marillac (Charente, Southwestern France): Morphology, comparisons and paleobiology
published pages: 102683, ISSN: 0047-2484, DOI: 10.1016/j.jhevol.2019.102683
Journal of Human Evolution 138 2020-01-28
2018 Figus C; Stephens NB; Sorrentino R; Panetta D; Belcastro MG; Ryan TM; Benazzi S
Unravelling morphological changes of the human talus during growth
published pages: 65, ISSN: , DOI:
Proceedings of the European Society for the study of Human Evolution 8 2020-01-28
2019 Carla Figus, Nicholas B Stephens, Eugenio Bortolini, Rita Sorrentino, Simona Arrighi, Federica Badino, Federico Lugli, Giulia Marciani, Gregorio Oxilia, Matteo Romandini, Lucia Martina Scalise, Sara Silvestrini, Maria Giovanna Belcastro, Timothy M Ryan, Stefano Benazzi
The ontogeny of bipedalism: insights from trabecular changes during growth.
published pages: 60, ISSN: , DOI:
Proceedings of the European Society for the study of Human Evolution 9 2020-01-28
2018 Lugli, Federico; Cipriani, Anna; Arnaud, Julie; Arzarello, Marta; Lembo, Giuseppe; Janssen, Renée; Vonhof, Hubert; Peretto, Carlo; Benazzi, Stefano
published pages: , ISSN: 2279-7335, DOI: 10.26382/aiqua.2018
Alpine and Mediterranean Quaternary 1 2019-12-16
2019 Jayde N. Hirniak, Eugene I. Smith, Racheal Johnsen, Minghua Ren, Jamie Hodgkins, Caley Orr, Fabio Negrino, Julien Riel‐Salvatore, Shelby Fitch, Christopher E. Miller, Andrea Zerboni, Guido S. Mariani, Jacob A. Harris, Claudine Gravel‐Miguel, David Strait, Marco Peresani, Stefano Benazzi, Curtis W. Marean
Discovery of cryptotephra at Middle–Upper Paleolithic sites Arma Veirana and Riparo Bombrini, Italy: a new link for broader geographic correlations
published pages: , ISSN: 0267-8179, DOI: 10.1002/jqs.3158
Journal of Quaternary Science 2019-12-16
2019 Matteo Romandini, Jacopo Crezzini, Eugenio Bortolini, Paolo Boscato, Francesco Boschin, Lisa Carrera, Nicola Nannini, Antonio Tagliacozzo, Gabriele Terlato, Simona Arrighi, Federica Badino, Carla Figus, Federico Lugli, Giulia Marciani, Gregorio Oxilia, Adriana Moroni, Fabio Negrino, Peresani Marco, Julien Riel-Salvatore, Annamaria Ronchitelli, Enza Elena Spinapolice, Stefano Benazzi
Macromammal and bird assemblages across the Late Middle to Upper Palaeolithic transition in Italy: an extended zooarchaeological review.
published pages: , ISSN: 1040-6182, DOI: 10.1016/j.quaint.2019.11.008
Quaternary International 2019-12-16
2019 Giulia Marciani, Annamaria Ronchitelli, Simona Arrighi, Federica Badino, Eugenio Bortolini, Paolo Boscato, Francesco Boschin, Jacopo Crezzini, Davide Delpiano, Armando Falcucci, Carla Figus, Federico Lugli, Gregorio Oxilia, Matteo Romandini, Julien Riel-Salvatore, Fabio Negrino, Marco Peresani, Enza Elena Spinapolice, Adriana Moroni, Stefano Benazzi
Lithic techno-complexes in Italy from 50 to 39 thousand years BP: An overview of lithic technological changes across the Middle-Upper Palaeolithic boundary
published pages: 27, ISSN: 1040-6182, DOI: 10.1016/j.quaint.2019.11.005
Quaternary International 2019-12-16
2019 Marco Peresani, Stefano Bertola, Davide Delpiano, Stefano Benazzi, Matteo Romandini
The Uluzzian in the north of Italy: insights around the new evidence at Riparo Broion
published pages: 3503-3536, ISSN: 1866-9557, DOI: 10.1007/s12520-018-0770-z
Archaeological and Anthropological Sciences 11/7 2019-11-28
2019 Adriana Moroni, Giovanni Boschian, Jacopo Crezzini, Guido Montanari-Canini, Giulia Marciani, Giulia Capecchi, Simona Arrighi, Daniele Aureli, Claudio Berto, Margherita Freguglia, Astolfo Araujo, Sem Scaramucci, Jean Jacques Hublin, Tobias Lauer, Stefano Benazzi, Fabio Parenti, Marzia Bonato, Stefano Ricci, Sahra Talamo, Aldo G. Segre, Francesco Boschin, Vincenzo Spagnolo
Late Neandertals in central Italy. High-resolution chronicles from Grotta dei Santi (Monte Argentario - Tuscany)
published pages: 130-151, ISSN: 0277-3791, DOI: 10.1016/j.quascirev.2018.11.021
Quaternary Science Reviews 217 2019-11-28
2019 Yohannes Haile-Selassie, Stephanie M. Melillo, Antonino Vazzana, Stefano Benazzi, Timothy M. Ryan
A 3.8-million-year-old hominin cranium from Woranso-Mille, Ethiopia
published pages: 214-219, ISSN: 0028-0836, DOI: 10.1038/s41586-019-1513-8
Nature 573/7773 2019-11-28
2019 Federica Badino, Roberta Pini, Cesare Ravazzi, Davide Margaritora, Simona Arrighi, Eugenio Bortolini, Carla Figus, Biagio Giaccio, Federico Lugli, Giulia Marciani, Giovanni Monegato, Adriana Moroni, Fabio Negrino, Gregorio Oxilia, Marco Peresani, Matteo Romandini, Annamaria Ronchitelli, Enza E. Spinapolice, Andrea Zerboni, Stefano Benazzi
An overview of Alpine and Mediterranean palaeogeography, terrestrial ecosystems and climate history during MIS 3 with focus on the Middle to Upper Palaeolithic transition
published pages: , ISSN: 1040-6182, DOI: 10.1016/j.quaint.2019.09.024
Quaternary International 2019-11-28
2019 Federico Lugli, Giulia Di Rocco, Antonino Vazzana, Filippo Genovese, Diego Pinetti, Elisabetta Cilli, Maria Cristina Carile, Sara Silvestrini, Gaia Gabanini, Simona Arrighi, Laura Buti, Eugenio Bortolini, Anna Cipriani, Carla Figus, Giulia Marciani, Gregorio Oxilia, Matteo Romandini, Rita Sorrentino, Marco Sola, Stefano Benazzi
Enamel peptides reveal the sex of the Late Antique ‘Lovers of Modena’
published pages: , ISSN: 2045-2322, DOI: 10.1038/s41598-019-49562-7
Scientific Reports 9/1 2019-11-28
2019 Katsuhiro Sano, Simona Arrighi, Chiaramaria Stani, Daniele Aureli, Francesco Boschin, Ivana Fiore, Vincenzo Spagnolo, Stefano Ricci, Jacopo Crezzini, Paolo Boscato, Monica Gala, Antonio Tagliacozzo, Giovanni Birarda, Lisa Vaccari, Annamaria Ronchitelli, Adriana Moroni, Stefano Benazzi
The earliest evidence for mechanically delivered projectile weapons in Europe
published pages: 1409-1414, ISSN: 2397-334X, DOI: 10.1038/s41559-019-0990-3
Nature Ecology & Evolution 3/10 2019-11-28
2019 Luca Fiorenza, Stefano Benazzi, Ottmar Kullmer, Giulia Zampirolo, Arnaud Mazurier, Clément Zanolli, Roberto Macchiarelli
Dental macrowear and cortical bone distribution of the Neanderthal mandible from Regourdou (Dordogne, Southwestern France)
published pages: 174-188, ISSN: 0047-2484, DOI: 10.1016/j.jhevol.2019.05.005
Journal of Human Evolution 132 2019-11-28
2019 Federico Lugli, Anna Cipriani, Giulia Capecchi, Stefano Ricci, Francesco Boschin, Paolo Boscato, Paola Iacumin, Federica Badino, Marcello A. Mannino, Sahra Talamo, Michael P. Richards, Stefano Benazzi, Annamaria Ronchitelli
Strontium and stable isotope evidence of human mobility strategies across the Last Glacial Maximum in southern Italy
published pages: 905-911, ISSN: 2397-334X, DOI: 10.1038/s41559-019-0900-8
Nature Ecology & Evolution 3/6 2019-11-28
2018 Matteo Romandini, Gabriele Terlato, Nicola Nannini, Antonio Tagliacozzo, Stefano Benazzi, Marco Peresani
Bears and humans, a Neanderthal tale. Reconstructing uncommon behaviors from zooarchaeological evidence in southern Europe
published pages: 71-91, ISSN: 0305-4403, DOI: 10.1016/j.jas.2017.12.004
Journal of Archaeological Science 90 2019-05-06
2017 Federico Lugli, Anna Cipriani, Julie Arnaud, Marta Arzarello, Carlo Peretto, Stefano Benazzi
Suspected limited mobility of a Middle Pleistocene woman from Southern Italy: strontium isotopes of a human deciduous tooth
published pages: , ISSN: 2045-2322, DOI: 10.1038/s41598-017-09007-5
Scientific Reports 7/1 2019-05-06
2018 Gregorio Oxilia, Eugenio Bortolini, Sergio Martini, Andrea Papini, Marco Boggioni, Laura Buti, Carla Figus, Rita Sorrentino, Grant Townsend, John Kaidonis, Luca Fiorenza, Emanuela Cristiani, Ottmar Kullmer, Jacopo Moggi‐Cecchi, Stefano Benazzi
The physiological linkage between molar inclination and dental macrowear pattern
published pages: , ISSN: 0002-9483, DOI: 10.1002/(ISSN)1096-8644
American Journal of Physical Anthropology 2019-05-06
2018 Federico Lugli, Anna Cipriani, Valentina Tavaglione, Mirko Traversari, Stefano Benazzi
Transhumance pastoralism of Roccapelago (Modena, Italy) early-modern individuals: Inferences from Sr isotopes of hair strands
published pages: 470-483, ISSN: 0002-9483, DOI: 10.1002/ajpa.23643
American Journal of Physical Anthropology 167/3 2019-05-06
2018 Adriana Moroni, Annamaria Ronchitelli, Simona Arrighi, Daniele Aureli, Shara E. Bailey, Paolo Boscato, Francesco Boschin, Giulia Capecchi, Jacopo Crezzini, Katerina Douka, Giulia Marciani, Daniele Panetta, Filomena Ranaldo, Stefano Ricci, Sem Scaramucci, Vincenzo Spagnolo, Stefano Benazzi, Paolo Gambassini
Grotta del Cavallo (Apulia – Southern Italy). The Uluzzian in the mirror
published pages: , ISSN: 2037-0644, DOI: 10.4436/JASS.96004
Journal of Anthropological Sciences 2019-05-06
2017 Laura Buti, Adeline Le Cabec, Daniele Panetta, Maria Tripodi, Piero A. Salvadori, Jean-Jacques Hublin, Robin N.M. Feeney, Stefano Benazzi
3D enamel thickness in Neandertal and modern human permanent canines
published pages: 162-172, ISSN: 0047-2484, DOI: 10.1016/j.jhevol.2017.08.009
Journal of Human Evolution 113 2019-05-06
2017 Cristiana Margherita, Gregorio Oxilia, Veronica Barbi, Daniele Panetta, Jean-Jacques Hublin, David Lordkipanidze, Tengiz Meshveliani, Nino Jakeli, Zinovi Matskevich, Ofer Bar-Yosef, Anna Belfer-Cohen, Ron Pinhasi, Stefano Benazzi
Morphological description and morphometric analyses of the Upper Palaeolithic human remains from Dzudzuana and Satsurblia caves, western Georgia
published pages: 83-90, ISSN: 0047-2484, DOI: 10.1016/j.jhevol.2017.07.011
Journal of Human Evolution 113 2019-05-06
2018 Rita Sorrentino, Eugenio Bortolini, Federico Lugli, Giuseppe Mancuso, Laura Buti, Gregorio Oxilia, Antonino Vazzana, Carla Figus, Maria Cristina Serrangeli, Cristiana Margherita, Annachiara Penzo, Giorgio Gruppioni, Antonio Gottarelli, Klaus Peter Jochum, Maria Giovanna Belcastro, Anna Cipriani, Robin N. M. Feeney, Stefano Benazzi
Unravelling biocultural population structure in 4th/3rd century BC Monterenzio Vecchio (Bologna, Italy) through a comparative analysis of strontium isotopes, non-metric dental evidence, and funerary practices
published pages: e0193796, ISSN: 1932-6203, DOI: 10.1371/journal.pone.0193796
PLOS ONE 13/3 2019-05-06

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